Sea turtles from the Upper Cretaceous of the Penza Province (Russia)

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The article describes remains of sea turtles (clade Pan-Chelonioidea) from three Upper Cretaceous localities of the Penza Province of Russia: Malaya Serdoba 1 and 2 (Campanian) and Penza (Maastrichtian). The distal part of the humerus of Pan-Chelonioidea indet. from the Malaya Serdoba 1 locality is similar to the humerus of Terlinguachelys fischbecki (Protostegidae) from the Campanian of USA. The turtle assemblage from the Malaya Serdoba 2 locality includes Protostega gigas (Protostegidae) and Pan-Chelonioidea indet. similar to T. fischbecki and representatives of the clade Ctenochelyidae, and resembles in systematic composition the sea turtle assemblage from the Campanian Beloe Ozero locality, Saratov Province of Russia. The outer surface of the costal plates of turtle from the Penza locality has a sculpture similar to Peritresius ornatus (Ctenochelyidae) and Glyptochelone suyckerbuycki.

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作者简介

E. Zvonok

Lugansk State Pedagogical University

Email: turtle@zin.ru
俄罗斯联邦, Lugansk, 291011

M. Arkhangelsky

Saratov State University; Saratov State Technical University

Email: turtle@zin.ru
俄罗斯联邦, Saratov, 410012; Saratov, 410054

Yu. Glazunov

Email: turtle@zin.ru

Independent researcher

俄罗斯联邦

V. Glazunov

Email: turtle@zin.ru

Independent researcher

俄罗斯联邦

A. Glushkov

Email: turtle@zin.ru

Independent researcher

俄罗斯联邦

I. Danilov

Zoological Institute, Russian Academy of Sciences

编辑信件的主要联系方式.
Email: turtle@zin.ru
俄罗斯联邦, St. Petersburg, 199034

参考

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2. Fig. 1. The Malaya Serdoba 2 locality: geographical position (A–B) and stratigraphic section (G). Designations: 1 – sand, 2 – sandstone, 3 – clay, 4 – siltstone, 5 – marl, 6 – opoka, 7 – phosphorites, 8 – glauconite, 9 – ferrugination, 10 – remains of reptiles, 11 – remains of sharks, 12 – remains of bony fish, 13 – traces of bioturbation, 14 – location and level of finds.

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3. Fig. 2. Fragments of two costal plates in the articulation of Pan-Chelonioidea indet., specimen ZIN PH 1/245, from above, from the Maastrichtian of Penza (a, b), and their labels (c–d): a – photograph; b – explanatory drawing; c – label by M.A. Vedenyapin; d – label by L.A. Nesov. The white line shows the approximate outlines of the costal ridges on the ventral surface of the plates. The dashed black line shows the reconstructed outlines of the free ribs. Scale bar – 3 cm.

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4. Fig. 3. Left maxilla of Protostega gigas Cope, 1872, specimen ZIN PH 1/291: a – lateral view; b – ventral view; c – anterior view; g, d – medial view (d – explanatory drawing); Penza region, Serdobsky district, Malaya Serdoba 2; Upper Cretaceous, Campanian, Rybushkinskaya suite. Key: ane – apertura narium externa; ani – apertura narium interna; da – depression lateral to apertura narium interna; fas – foramen of alveolare superius; fn – fossa nasalis; fsm – (?) foramen supramaxillare; gas – groove leading to foramen of alveolare superius; lar – labial ridge; lir – lingual ridge; mg – medial groove; o – orbit; sj – sutural surface for the zygomatic bone; spf – sutural surface for the prefrontal bone; spl – sutural surface for the palatine bone; spm – sutural surface for the premaxilla; sv – (?) worn sutural surface for the vomer. White line – edge of the suture for the premaxilla on the outer surface of the maxilla, dashed black line – reconstructed edges of the bone. Scale bar – 5 cm.

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5. Fig. 4. Bones of Pan-Chelonioidea indet.: a–d – right humerus, specimen ZIN PH 2/291: a, b – ventral view; c, g – dorsal view; d – anterior view; f–i – right femur, specimen ZIN PH 3/291: f, g – ventral view; h – anterior view; i – view from the proximal end; k–m – right V or VI costal plate, specimen ZIN PH 4/291: k – ventral view; l – view from the medial edge; m – dorsal view; n–p – left VIII costal plate, specimen ZIN PH 5/291: n – dorsal view; o – view from the medial edge; p – ventral view; p–y – right (?) VIII peripheral plate, specimen. ZIN PH 6/291: p – dorsal view; c – ventral view; t – anterior view, y – posterior view; Penza region, Serdobsky district, Malaya Serdoba 2; Upper Cretaceous, Campanian, Rybushkinskaya suite. Key: cf – head of the femur; ch – head of the humerus; dcb – depression for attachment of mm. coracobrachialis brevis; dld – (?) depression for attachment of mm. latissimus dorsi et teres major; lp – lateral process; ims – intermarginal groove; mat – greater trochanter; mit – lesser trochanter; mp – medial process; rIX – IX rib; rX – articular surface for the X rib. Scale bar – 3 cm.

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6. Fig. 5. Distal part of the right humerus of Pan-Chelonioidea indet., specimen SGU 104a/32: a – dorsal view; b – posterior and ventral views; c – posterior view, g – anterior view, d – view from the distal end; Penza Region, Serdobsky District, Malaya Serdoba 1; Upper Cretaceous, Campanian, Rybushkinskaya Formation. Designations: c – capitellum; eef – ectepicondylar foramen; t – block. Scale bar – 5 cm.

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